Data Zone
Barlow, K. E., Boyd, I. L., Croxall, J. P., Reid, K., Staniland, I. J.& Brierley, A. S. 2002. Are penguins and seals in competition for Antarctic krill at South Georgia? Marine Biology 140: 205-213.
Barlow, K.E.; Croxall, J.P. 2002. Seasonal and interannual variation in foraging range and habitat of macaroni penguins at South Georgia. Marine Ecology Progress Series 232: 291–304.
Bird, J.P., Martin, R., Akçakaya, H.R., Gilroy, J., Burfield, I.J., Garnett, S.G., Symes, A., Taylor, J., Şekercioğlu, Ç.H. and Butchart, S.H.M. 2020. Generation lengths of the world’s birds and their implications for extinction risk. Conservation Biology 34(5): 1252-1261.
Bon, C., Penna, A.D., D’Ovidio, F., Arnould,J.P.Y., Bost, C.A. 2015. Influence of oceanographic structures on foraging strategies: Macaroni penguins at Crozet. Movement Ecology 3: 32, DOI 10.1186/s40462-015-0057-2.
Bost, C. A.; Thiebot, J. B.; Pinaud, D.; Cherel, Y.; Trathan, P. N. 2009. Where do penguins go during the inter-breeding period? Using geolocation to track the winter dispersion of the Macaroni Penguin. Biology Letters 5: 473-476.
Cooper, J.; Crawford, R. J. M.; De Villiers, M. S.; Dyer, B. M.; Hofmeyr, G. J. G.; Jonker, A. 2009. Disease outbreaks among penguins at sub-Antarctic Marion Island: A conservation concern. Marine Ornithology 37: 193–196.
Crawford, R. J. M., Whittington, P. A., Upfold, L., Ryan, P. G., Petersen, S. L., Dyer, B. M. & Cooper, J. 2009. Recent trends in numbers of four species of penguins at the Prince Edward Islands. African Journal of Marine Science 31(3): 419-426.
Crossin, G. T., Trathan, P. and Crawford, R. J. M. 2013. Macaroni Penguin (Eudyptes chrysolophus) and Royal Penguin (Eudyptes schlegeli). In: P. G. Borboroglu and P. D. Boersma (eds), Penguins: Natural History and Conservation, pp. 185-208. University of Washington Press, Seattle.
Crossin, G.T.; Takahashi, A,; Sakamoto, K.Q.; Trathan, P.N.; Williams, T.D. 2015. Habitat selection by foraging macaroni penguins correlates with hematocrit, an index of aerobic condition. Marine Ecology Progress Series 530: 163-176.
Crowley, G. M. and Garnett, S. T. 2000. Changing Fire Management in the Pastoral Lands of Cape York Peninsula of northeast Australia, 1623 to 1996. Australian Geographical Studies 38(1): 10–26.
Deagle, B. E., Gales, N. J. & Hindell, M. A. 2008. Variability in foraging behaviour of chick-rearing macaroni penguins Eudyptes chrysolophus and its relation to diet. Marine Ecology Progress Series 359: 295-309.
Dyer, B.M; Crawford, R.J.M. 2015. Macaroni penguin. In: Taylor, M.R.; Peacock, F.; Wanless, R.M. The 2015 ESKOM Red Data Book of Birds. BirdLife International..
Ellis, S., Croxall, J. P. & Cooper, J. 1998. Penguin conservation assessment and management plan: report from the workshop held 8-9 September 1996, Cape Town, South Africa. IUCN/SSC, Apple Valley, USA.
Frugone, M. J., López, M. E., Segovia, N. I., Cole, T. L., Lowther, A., Pistorius, P., Dantas, G. P., Petry, M. V., Bonadonna, F., Trathan, P. & Polanowski, A. 2019. More than the eye can see: Genomic insights into the drivers of genetic differentiation in Royal/Macaroni penguins across the Southern Ocean. Molecular phylogenetics and evolution 139: 106563.
Frugone, M. J., Lowther, A., Noll, D., Ramos, B., Pistorius, P., Dantas, G. P. M., Petry, M. V., Bonadonna, F., Steinfurth, A., Polanowski, A. & Rey, A. R. 2018. Contrasting phylogeographic pattern among Eudyptes penguins around the Southern Ocean. Scientific reports 8(1): 1-16.
Horswill, C., Matthiopoulos, J., Green, J. A., Meredith, M. P., Forcada, J., Peat, H., Preston, M., Trathan, P.N. and Ratcliffe, N. 2014. Survival in macaroni penguins and the relative importance of different drivers: individual traits, predation pressure and environmental variability. The Journal of Animal Ecology 83(5): 1057–1067.
Horswill, C., Ratcliffe, N., Green, J. A., Phillips, R. A., Trathan, P. N., & Matthiopoulos, J. 2016. Unravelling the relative roles of top‐down and bottom‐up forces driving population change in an oceanic predator. Ecology 97(8): 1919–1928.
Isaksen, K., Hofmeyr, G.J.G., Dyer, B.M.,Naestvold, A., Mehlum, F., Gjertz, I., Bakken, V., Huyser, O. 1997. Preliminary results from CEMP-monitoring of Antarctic Fur Seals, Chinstrap Penguins and Macaroni Penguins at Bouvetoya 1996/97. Report to CCAMLR WG-EMM 1997: 26.
IUCN. 2020. The IUCN Red List of Threatened Species. Version 2020-3. Available at: https://www.iucnredlist.org. (Accessed: 10 December 2020).
Kirkwood, R.; Lawton, K.; Moreno, C.; Valencia, J.; Schlatter, R.; Robertson, G. 2007. Estimates of southern rockhopper and macaroni penguin numbers at the Ildefonso and Diego Ramirez archipelagos, Chile, using quadrat and distance-sampling techniques. Waterbirds 30 : 259-267.
Marchant, S.; Higgins, P. J. 1990. Handbook of Australian, New Zealand and Antarctic birds, 1: ratites to ducks. Oxford University Press, Melbourne.
Oehler, D.A.; Pelikan, S., Fry, W.R.; Weakley, L.; Kush, A.; Marin, M. 2008. Status of crested penguins Eudyptes spp. on three islands in southern Chile. Wilson. Wilson Journal of Ornithology 120: 575–81.
Ratcliffe, N.; Hill, S.L.; Staniland, I.J.; Brown, R.; Adlard, S.; Horswill, C.; Trathan, P.N. 2015. Do krill fisheries compete with macaroni penguins? Spatial overlap in prey consumption and catches during winter. Diversity and Distributions 21: 1339-1348.
Thiebot, J.B.; Cherel, Y.; Trathan, P.N.; Bost, C.A. 2011. Inter-population segregation and in the wintering areas of macaroni penguins. Marine Ecology Progress Series 421 : 279–290.
Trathan, P. N.; Croxall, J. P.; Murphy, E. J.; Everson, I. 1998. Use of at-sea distribution data to derive potential foraging ranges of macaroni penguins during the breeding season. Marine Ecology Progress Series 169: 263-275.
Trathan, P.N., Garcia-Borboroglu, P., Boersma, D., Bost, C.A., Crawford, R.J.M., Crossin, G.T., Cuthbert, R.J., Dann, P., Davis, L.S., De La Puente, S., Ellenberg, U., Lynch, H.J., Mattern, T., Putz, K., Seddon, P.J., Trivelpiece, W. and Wienecke, B. 2014. Pollution, habitat loss, fishing, and climate change as critical threats to penguins. Conservation Biology 29(1): 31-41.
Trathan, P.N., Green, C., Tanton, J., Peat, H., Poncet, J., Morton A. 2006. Foraging dynamics of macaroni penguins Eudyptes chryolophus at South Georgia during brood-guard. Marine Ecology-Progress Series 323: 239-251.
Trathan, P.N.; Collins, M.A.; Grant, S.M.; Belchier, M.; Barnes, D.K.A.; Brown, J.; Staniland, I.J. 2014. The South Georgia and the South Sandwich Islands MPA: Protecting a biodiverse oceanic island chain situated in the flow of the Antarctic Circumpolar Current. In: Johnson, M.L.; Sandell, J. (ed.), Marine Managed Areas and Fisheries: Advances in Marine Biology 69: 15-78..
Trathan, P.N.; Ratcliffe, N.; Masden, E.A. 2012. Ecological drivers of change at South Georgia: the krill surplus, or climate variability. Ecography 35: 983-993.
Whitehead T.O., Kato, A., Ropert-Coudert Y, Ryan, P. 2016. Habitat use and diving behaviour of macaroni Eudyptes chrysolophus and eastern rockhopper E. chrysocome filholi penguins during the critical pre-moult period. Marine Biology 163: 19-37.
Woehler, E.J. 1993. The distribution and abundance of Antarctic and Subantarctic penguins. Scientific Commission on Antarctic Research, Cambridge, U.K.
IUCN SSC Penguin Specialist Group
Australian Govt - Action Plan for Australian Birds 2000 - Recovery Outline
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Recommended citation
BirdLife International (2024) Species factsheet: Macaroni Penguin Eudyptes chrysolophus. Downloaded from
https://datazone.birdlife.org/species/factsheet/macaroni-penguin-eudyptes-chrysolophus on 23/11/2024.
Recommended citation for factsheets for more than one species: BirdLife International (2024) IUCN Red List for birds. Downloaded from
https://datazone.birdlife.org/species/search on 23/11/2024.