Macaroni Penguin Eudyptes chrysolophus


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Barlow, K.E.; Croxall, J.P. 2002. Seasonal and interannual variation in foraging range and habitat of macaroni penguins at South Georgia. Marine Ecology Progress Series 232: 291–304.

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Bon, C., Penna, A.D., D’Ovidio, F., Arnould,J.P.Y., Bost, C.A. 2015. Influence of oceanographic structures on foraging strategies: Macaroni penguins at Crozet. Movement Ecology 3: 32, DOI 10.1186/s40462-015-0057-2.

Bost, C. A.; Thiebot, J. B.; Pinaud, D.; Cherel, Y.; Trathan, P. N. 2009. Where do penguins go during the inter-breeding period? Using geolocation to track the winter dispersion of the Macaroni Penguin. Biology Letters 5: 473-476.

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Crossin, G. T., Trathan, P. and Crawford, R. J. M. 2013. Macaroni Penguin (Eudyptes chrysolophus) and Royal Penguin (Eudyptes schlegeli). In: P. G. Borboroglu and P. D. Boersma (eds), Penguins: Natural History and Conservation, pp. 185-208. University of Washington Press, Seattle.

Crossin, G.T.; Takahashi, A,; Sakamoto, K.Q.; Trathan, P.N.; Williams, T.D. 2015. Habitat selection by foraging macaroni penguins correlates with hematocrit, an index of aerobic condition. Marine Ecology Progress Series 530: 163-176.

Crowley, G. M. and Garnett, S. T. 2000. Changing Fire Management in the Pastoral Lands of Cape York Peninsula of northeast Australia, 1623 to 1996. Australian Geographical Studies 38(1): 10–26.

Deagle, B. E., Gales, N. J. & Hindell, M. A. 2008. Variability in foraging behaviour of chick-rearing macaroni penguins Eudyptes chrysolophus and its relation to diet. Marine Ecology Progress Series 359: 295-309.

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Frugone, M. J., López, M. E., Segovia, N. I., Cole, T. L., Lowther, A., Pistorius, P., Dantas, G. P., Petry, M. V., Bonadonna, F., Trathan, P. & Polanowski, A. 2019. More than the eye can see: Genomic insights into the drivers of genetic differentiation in Royal/Macaroni penguins across the Southern Ocean. Molecular phylogenetics and evolution 139: 106563.

Frugone, M. J., Lowther, A., Noll, D., Ramos, B., Pistorius, P., Dantas, G. P. M., Petry, M. V., Bonadonna, F., Steinfurth, A., Polanowski, A. & Rey, A. R. 2018. Contrasting phylogeographic pattern among Eudyptes penguins around the Southern Ocean. Scientific reports 8(1): 1-16.

Horswill, C., Matthiopoulos, J., Green, J. A., Meredith, M. P., Forcada, J., Peat, H., Preston, M., Trathan, P.N. and Ratcliffe, N. 2014. Survival in macaroni penguins and the relative importance of different drivers: individual traits, predation pressure and environmental variability. The Journal of Animal Ecology 83(5): 1057–1067.

Horswill, C., Ratcliffe, N., Green, J. A., Phillips, R. A., Trathan, P. N., & Matthiopoulos, J. 2016. Unravelling the relative roles of top‐down and bottom‐up forces driving population change in an oceanic predator. Ecology 97(8): 1919–1928.

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Ratcliffe, N.; Hill, S.L.; Staniland, I.J.; Brown, R.; Adlard, S.; Horswill, C.; Trathan, P.N. 2015. Do krill fisheries compete with macaroni penguins? Spatial overlap in prey consumption and catches during winter. Diversity and Distributions 21: 1339-1348.

Thiebot, J.B.; Cherel, Y.; Trathan, P.N.; Bost, C.A. 2011. Inter-population segregation and in the wintering areas of macaroni penguins. Marine Ecology Progress Series 421 : 279–290.

Trathan, P. N.; Croxall, J. P.; Murphy, E. J.; Everson, I. 1998. Use of at-sea distribution data to derive potential foraging ranges of macaroni penguins during the breeding season. Marine Ecology Progress Series 169: 263-275.

Trathan, P.N., Garcia-Borboroglu, P., Boersma, D., Bost, C.A., Crawford, R.J.M., Crossin, G.T., Cuthbert, R.J., Dann, P., Davis, L.S., De La Puente, S., Ellenberg, U., Lynch, H.J., Mattern, T., Putz, K., Seddon, P.J., Trivelpiece, W. and Wienecke, B. 2014. Pollution, habitat loss, fishing, and climate change as critical threats to penguins. Conservation Biology 29(1): 31-41.

Trathan, P.N., Green, C., Tanton, J., Peat, H., Poncet, J., Morton A. 2006. Foraging dynamics of macaroni penguins Eudyptes chryolophus at South Georgia during brood-guard. Marine Ecology-Progress Series 323: 239-251.

Trathan, P.N.; Collins, M.A.; Grant, S.M.; Belchier, M.; Barnes, D.K.A.; Brown, J.; Staniland, I.J. 2014. The South Georgia and the South Sandwich Islands MPA: Protecting a biodiverse oceanic island chain situated in the flow of the Antarctic Circumpolar Current. In: Johnson, M.L.; Sandell, J. (ed.), Marine Managed Areas and Fisheries: Advances in Marine Biology 69: 15-78..

Trathan, P.N.; Ratcliffe, N.; Masden, E.A. 2012. Ecological drivers of change at South Georgia: the krill surplus, or climate variability. Ecography 35: 983-993.

Whitehead T.O., Kato, A., Ropert-Coudert Y, Ryan, P. 2016. Habitat use and diving behaviour of macaroni Eudyptes chrysolophus and eastern rockhopper E. chrysocome filholi penguins during the critical pre-moult period. Marine Biology 163: 19-37.

Woehler, E.J. 1993. The distribution and abundance of Antarctic and Subantarctic penguins. Scientific Commission on Antarctic Research, Cambridge, U.K.

Further resources

IUCN SSC Penguin Specialist Group

Australian Govt - Action Plan for Australian Birds 2000 - Recovery Outline

Search for photos and videos, and hear sounds of this species from the Macaulay Library

Media resources

Recommended citation
BirdLife International (2022) Species factsheet: Eudyptes chrysolophus. Downloaded from on 08/12/2022. Recommended citation for factsheets for more than one species: BirdLife International (2022) IUCN Red List for birds. Downloaded from on 08/12/2022.