Corncrake Crex crex


Taxonomic source(s)
AERC TAC. 2003. AERC TAC Checklist of bird taxa occurring in Western Palearctic region, 15th Draft. Available at: # _the_WP15.xls#.
AERC TAC. 2003. AERC TAC Checklist of bird taxa occurring in Western Palearctic region, 15th Draft. Available at: # _the_WP15.xls#.
Christidis, L. and Boles, W.E. 2008. Systematics and taxonomy of Australian birds. CSIRO Publishing, Collingwood, Australia.
Christidis, L.; Boles, W. E. 2008. Systematics and taxonomy of Australian birds. CSIRO Publishing, Collingwood, Australia.
Cramp, S. and Perrins, C.M. 1977-1994. Handbook of the birds of Europe, the Middle East and Africa. The birds of the western Palearctic. Oxford University Press, Oxford.
Cramp, S.; Perrins, C. M. 1977-1994. Handbook of the birds of Europe, the Middle East and Africa. The birds of the western Palearctic. Oxford University Press, Oxford.
del Hoyo, J., Collar, N.J., Christie, D.A., Elliott, A. and Fishpool, L.D.C. 2014. HBW and BirdLife International Illustrated Checklist of the Birds of the World. Lynx Edicions BirdLife International, Barcelona, Spain and Cambridge, UK.
del Hoyo, J.; Collar, N. J.; Christie, D. A.; Elliott, A.; Fishpool, L. D. C. 2014. HBW and BirdLife International Illustrated Checklist of the Birds of the World. Barcelona, Spain and Cambridge UK: Lynx Edicions and BirdLife International.

IUCN Red list criteria met and history
Red List criteria met
Critically Endangered Endangered Vulnerable
- - -

Red List history
Year Category Criteria
2016 Least Concern
2012 Least Concern
2010 Least Concern
2008 Near Threatened A3c
2006 Near Threatened
2004 Near Threatened
2000 Vulnerable
1996 Vulnerable
1994 Vulnerable
1988 Threatened
Species attributes

Migratory status full migrant Forest dependency Does not normally occur in forest
Land mass type Land-mass type - continent
Average mass -

Estimate Data quality
Extent of Occurrence breeding/resident (km2) 20,600,000 medium
Extent of Occurrence non-breeding (km2) 7,070,000 medium
Number of locations 11-100 -
Severely Fragmented -
Population and trend
Value Data quality Derivation Year of estimate
No. of mature individuals 3000000-7000000 medium estimated 2012
Population trend Stable poor estimated -
Decline (3 years/1 generation past) - - -
Decline (5 years/1 generation past) - - -
Decline (10 years/1 generation past) - - -
Decline (10 years/3 generation future) 1-19 - - -
Decline (10 years/3 generation past and future) - - -
Number of subpopulations 2-100 - - -
Percentage in largest subpopulation 1-89 - - -
Generation length (yrs) 3.7 - - -

Population justification: The European population is estimated at 1,290,000-2,120,000 calling or lekking males, which equates to 2,590,000-4,240,000 mature individuals (including 1-1.5 million pairs in European Russia) (BirdLife International 2015). This is marginally higher than the last European population estimate of 1.3-2 million breeding pairs (BirdLife International 2004) and higher than the 1.1-1.8 million pairs previously estimated by Schäffer and Mammen (1999). A further 515,000-1,240,000 pairs are estimated for Asiatic Russia, yielding a global total of 3,600,000-6,700,000 mature individuals. Given the high level of uncertainty in some of the breeding estimates and the apparent scarcity of the species in its non-breeding areas in sub-Saharan Africa, the total population may fall at the lower end of this range; even in the low millions.

Trend justification: Ongoing monitoring since 2002 in Russia (which holds the vast majority of the global population) indicates that numbers have remained stable or are even increasing, with some fluctuations due to extreme weather (A. Mischenko in litt. 2010). Whilst it is difficult to accurately predict future trends owing to the species's extensive range and differing climatic and agricultural conditions in different regions, it is thought that populations in key parts of the range in Russia and Kazakhstan are unlikely to change dramatically in the near future. Its conservation status remains unfavourable in some parts of its range with declines reported in several range countries (BirdLife International Corncrake Conservation Team 2015) although the overall European population trend was recently estimated as stable (BirdLife International 2015). Land-use changes are precautionarily predicted to drive a future decline of 1-19% in the forthcoming three generation (11 year) period.

Country/territory distribution
Country/Territory Occurrence status Presence Resident Breeding Non-breeding Passage
Afghanistan V Extant
Albania N Extant Yes Yes
Algeria N Extant Yes
Angola V Extant Yes
Armenia N Extant Yes Yes
Austria N Extant Yes
Azerbaijan N Extant Yes Yes
Bahrain N Extant Yes
Belarus N Extant Yes
Belgium N Extant Yes
Bermuda (to UK) V Extant Yes
Bosnia and Herzegovina N Extant Yes
Botswana V Extant Yes
Bulgaria N Extant Yes Yes
Cameroon V Extant Yes
Chad V Extant Yes
China (mainland) N Extant Yes
Congo N Extant Yes
Congo, The Democratic Republic of the N Extant Yes
Côte d'Ivoire V Extant Yes
Croatia N Extant Yes
Cyprus N Extant Yes Yes
Czechia N Extant Yes
Denmark N Extant Yes Yes
Egypt N Extant Yes
Eritrea V Extant Yes
Estonia N Extant Yes
Eswatini N Extant
Ethiopia N Extant Yes
Faroe Islands (to Denmark) N Extinct Yes
Finland N Extant Yes
France N Extant Yes Yes
Gabon V Extant Yes
Georgia N Extant Yes Yes
Germany N Extant Yes
Ghana V Extant Yes
Greece N Extant Yes
Greenland (to Denmark) V Extant
Guinea U Extant Yes
Hungary N Extant Yes
Iceland V Extant
Iran, Islamic Republic of N Extant Yes
Iraq N Extant Yes
Ireland N Extant Yes
Israel N Extant Yes
Italy N Extant Yes Yes
Jordan N Extant Yes
Kazakhstan N Extant Yes
Kenya N Extant Yes Yes
Kuwait N Extant Yes
Kyrgyzstan N Extant Yes
Latvia N Extant Yes
Lebanon N Extant Yes
Lesotho V Extant
Libya V Extant
Liechtenstein N Extant Yes
Lithuania N Extant Yes
Luxembourg N Extant Yes
Malawi N Extant Yes
Mali V Extant Yes
Mauritania N Extant Yes
Moldova N Extant Yes Yes
Mongolia N Extant Yes
Montenegro N Extant Yes
Morocco N Extant Yes
Mozambique N Extant Yes
Namibia V Extant Yes
Netherlands N Extant Yes
Niger V Extant Yes
Nigeria V Extant Yes
North Macedonia N Extant Yes
Norway N Extant Yes
Oman N Extant Yes Yes
Palestine N Extant Yes
Poland N Extant Yes
Portugal N Extant Yes
Qatar N Extant Yes
Romania N Extant Yes Yes
Russia N Extant Yes Yes
Russia (Asian) N Extant Yes
Russia (Central Asian) N Extant Yes
Russia (European) N Extant Yes Yes
Rwanda V Extant Yes
Saudi Arabia N Extant Yes
Serbia N Extant Yes
Seychelles V Extant Yes
Slovakia N Extant Yes
Slovenia N Extant Yes
Somalia V Extant Yes
South Africa N Extant Yes
South Sudan N Extant Yes
Spain N Extant Yes
Sri Lanka N Extant Yes
St Pierre and Miquelon (to France) V Extant Yes
Sudan N Extant Yes Yes
Sweden N Extant Yes
Switzerland N Extant Yes
Syria N Extant Yes
Tajikistan N Extant Yes
Tanzania N Extant Yes
Tunisia N Extant Yes
Turkey N Extant Yes
Turkmenistan N Extant Yes
Uganda V Extant Yes
Ukraine N Extant Yes Yes
United Arab Emirates N Extant Yes
United Kingdom N Extant Yes
Uzbekistan N Extant Yes
Vietnam V Extant Yes
Yemen N Extant Yes
Zambia N Extant Yes
Zimbabwe N Extant Yes

Important Bird and Biodiversity Areas (IBA)
Country/Territory IBA Name
Czechia Trebonsko (Trebon region)
Czechia Sumava mountains (Bohemian forest)
Serbia Vršačke Planine
Serbia Labudovo okno
Serbia Deliblatska pescara
Serbia Carska bara
Bosnia and Herzegovina Bardaca
Azerbaijan Alty Agach area
Azerbaijan Kusari (Gusari) area
Azerbaijan Sheki upland
Belarus Yaselda
Belarus Sporovo
Belarus Balota Zvaniec
Belarus Mid Prypiac'
Belarus Lower Pripyat
Belarus Vyhanaščanskija baloty
Belarus Beresina
Belarus Bielaje fish farm
Belarus Sož floodplain
Belarus Halubickaja Pušča
Belarus Zachodniaja Biarezina
Belarus Biarezinski zapaviednik
Belarus Haradzienskaja Svislač
Belarus Al'manskija baloty
Lithuania Kretuonas Landscape Reserve
Lithuania Nemunas delta regional Park
Lithuania Zuvintas Biosphere Reserve
Lithuania Tyruliai State Nature Reserve
Lithuania Novaraistis State Ornithological Reserve
Lithuania Verkne river
Lithuania Valley of Sirvinta River
Lithuania Krekenava forests
Lithuania Sesuva river
Lithuania Kauno Marios water reservoir & eastern coastal zone
Lithuania Birzulius-Stervas marshes complex
Lithuania Islauzas fish-ponds
Lithuania Jura & Sesuvis rivers and the lower reaches of Jura River
Lithuania Sartai Regional Park and Rokiskis Fishponds
Liechtenstein Ruggeller Riet, Talgebiet
Estonia Matsalu Bay
Estonia Luitemaa
Estonia Põhja-Liivimaa
Estonia Soomaa
Estonia Puhatu mire
Estonia Mouth of the Emajõgi river and Piirissaar island
Estonia Alam-Pedja
Estonia Koiva-Mustjõe
Estonia Ropka-Ihaste
Cyprus Phasouri reedbeds
Latvia Slitere and Zilie kalni
Latvia Engure lake
Latvia Ziemelu bogs
Latvia Kamburi meadows
Latvia Pape
Latvia Liepaja lake
Latvia Sita and Pededze flood-plain
Latvia Aiviekste flood-plain
Latvia Uzava meadows
Latvia Zile forest area
Latvia Cirgali meadows
Latvia Berzpils bogs and flood-plains
Latvia Vidusburtnieks
Latvia Durbe lake meadows
Latvia Dviete flood-plain
Latvia Lielupe flood-plain meadows
Latvia Uzava lowland
Latvia Lubans and fish-ponds
Latvia Satini fish-ponds
Latvia Svete lower reaches
Latvia Ruja flood-plain
Latvia Kalnciems meadows and Odini fields
Bulgaria Atanasovsko Lake
Bulgaria Western Balkan
Bulgaria Orsoya Fish-ponds
Bulgaria Radinchevo
Bulgaria Strandzha
Bulgaria Pomorie Lake
Romania Satchinez marsh
Romania Stânca lake - Costeşti
Romania Miheşu de Câmpie-Taureni fish-ponds
Romania Danube Delta and Razelm-Sinoe complex
Romania Jijiei şi Miletinului fish-ponds
Romania Padiş - Vlădeasa mountain
Romania Balta Mică a Brăilei
Romania Comana
Romania Dobron forest
Romania Rarău Mountains - Giumalău
Ireland River Shannon callows: Portumna-Athlone
Ireland Inishbofin and Inishshark (including Davillaun)
Ireland Sheskinmore Lough
Ireland Tory Island
Ireland Inishbofin, Inisdooey and Inisbeg
Ireland Lough Conn and Lough Cullin (including Moy valley)
Ireland Fanad Head peninsula
Ireland Malin Head
Ireland Falcarragh to Min an Chladaigh
Ireland Broadhaven, Blacksod and Tullaghan Bays and parts of the Mullet peninsula
Slovenia Ljubljansko moor
Slovenia Lake Cerknica
Slovenia Planina polje
Slovenia Ribnica valley
Slovenia Nanoščica river basin
Slovenia Reka valley
Slovenia Julijci
Belgium Côte Bajocienne
Belgium Sinémurienne
Belgium Hautes Fagnes/Eifel
Belgium Marche en Famenne
Belgium Lesse et Lomme
Belgium Entre-Sambre-et-Meuse
Moldova Lower Prut River and Manta-Beleu Lake
Moldova Lower Dniester River
Moldova Beech Land Forest
Sweden River Dal – Håvran
Sweden River Svartån
Sweden Lake Hjälstaviken
Sweden Lake Kvismaren
Sweden Lake Östen
Sweden Lake Hornborga
Sweden Lake Tåkern
Sweden Getterön
Sweden Lake Åsnen
Sweden River Helge
Sweden River Klingaälv – Lake Kranke
Sweden Lake Ralången – Lake Hyllingen
Sweden Ledskär – Bay of Karlholm
Sweden Lake Tämnaren
Sweden Dannemora
Poland Biebrza river valley
Poland Bialowieza Forest
Poland Lower Bug river valley
Poland Liwiec river valley
Poland Middle Vistula River Valley
Poland Malopolska Wisla river gap
Poland Bubnow Marsh
Poland Chelm calcareous marshes
Poland Bieszczady Mountains
Poland Janów forests
Poland Tatra mountains
Poland Przemków ponds
Poland Warta River Mouth
Poland Notec river flood-plain
Poland Middle Warta River Valley
Poland Druzno lake
Poland Vistula Lagoon
Poland Slowinska Site
Poland Insko Site
Poland Lower Odra river valley
Poland Swidwie Site
Poland Kampinos forest
Poland Biala Forest
Poland Zegrzynski reservoir
Poland Dymerskie meadows
Poland Luknajno lake
Poland Oswin site
Poland Gródek-Michalowo basin
Poland Jeziorsko reservoir
Poland Delta of the Swina river
Poland Obra river flood-plain
Poland Tysmienica river valley
Poland Marshy Valley of the Narew River
Poland Upper Wisla River Valley
Poland Turawa reservoir
Poland Middle Bug river valley
Poland Nida Rivar Valley
Poland Ner River Valley
Poland Parczew forests
Poland Slesin and Wystep ponds
Poland Przemysl Plateau
Poland Pisz forest
Poland Miedwie site
Poland Marshy valley of the Drweca river
Poland Napiwodzko-Ramucka forest
Poland Borecka forest
Poland Nietlice marshes
Poland Augustów forest
Poland Oak-hornbeam forests in the Odra valley
Poland Upper Narew River Valley
Poland Rakutowskie swamps
Poland Slup reservoir
Poland Ilawa Forest
Poland Odra riverine forests
Poland Slupia River Valley
Poland Lower Vistula River Valley
Poland Omulew and Plodownica Rivers Valleys
Greece Kythira island
Greece Dionysades islets, Crete
Greece Lakes Chortarolimni and Alyki, Moudros gulf, Diapori fen and Fakos peninsula, Limnos
Netherlands IJssel: Deventer - Zwolle
Netherlands Rijn: Heteren - Amerongen
Netherlands Waal: Ewijk - Waardenburg
Netherlands Gelderse Poort
Netherlands Biesbosch
Finland Köyliönjärvi and Pyhäjärvi lakes
Finland Puurijärvi-Isosuo wetlands
Finland Pori archipelago and wetlands
Finland Hanko western archipelago
Finland Porvoonjoki Delta
Finland Teutjärvi and Suvijärvi lakes
Finland Kirkon-Vilkkiläntura Bay
Finland Kirkkojärvi lake and Lupinlahti bay
Finland Siikalahti bay and Sammallampi lake
Finland Torronsuo and Lake Talpianjärvi
Finland Koijärvi lake
Finland Artjärvi wetlands
Finland Kangasala wetlands
Finland Lake Pihlajavesi
Finland Maaninka wetlands
Finland Värtsilä valley
Finland Alajoki
Finland Lapväärti wetlands
Hungary Aggtelek Karst
Hungary Hortobágy and Tisza-to
Hungary Lake Kis-Balaton
Hungary Hanság
Hungary Bodrogzug–Taktaköz
Hungary Ipoly Valley
Hungary Nagy-berek of Fonyód
Hungary Kolon Lake
Hungary Kesznyéten
Hungary Szatmár-Bereg
Hungary Inner Somogy
Hungary Tápió valley
Russia (European) Delta of the River Don
Russia (European) Sura environs
Russia (European) Flood-plain of Khoper river near Il'men' lake
Russia (European) Sources of Osuga river
Russia (European) Smolenskoye Poozer'ye
Russia (European) North part of Volgogradski reservoir
Russia (European) Sources of the River Luga
Russia (European) Racheyski forest
Russia (European) Buzulukski forest
Russia (European) Lake Lacha
Russia (European) Zaonezh'ye
Russia (European) Kulaksay lowland
Russia (European) Watershed of Bityug and Tsna rivers
Russia (European) Zavoronezhski area
Russia (European) Valley of Sysola river
Russia (European) Olonets plain
Russia (European) Vorono-Khoperski area
Russia (European) Kargopol' area
Russia (European) Zavidovo Reserve, including Lotoshinski, Klinski and Diatlovo fish ponds
Russia (European) Faustovo flood-plains of Moscow river
Russia (European) Nerussa - Desna woodland
Russia (European) Nemanski forest
Russia (European) Caucasus Biosphere Reserve
Russia (European) Mordovian P. G. Smidovich Nature Reserve
Russia (European) Pereluchski Nature Reserve
Russia (European) Lake Il'men and adjoining marshy plain
Russia (European) Flood-plain of Volkhov river
Russia (European) Redrovski Nature Reserve
Russia (European) Polisto-Lovatskaya mire system
Russia (European) Chudsko-Pskovski Lake and the adjacent areas
Russia (European) Central Forest Biosphere Reserve and adjacent areas
Russia (European) Stakhovski marshes
Russia (European) Upper Mologa river (lake Verestovo)
Russia (European) Savtsinskoye marsh
Russia (European) Khoper Nature Reserve
Russia (European) Voronezhski Nature Reserve
Russia (European) Khrenovski forest
Russia (European) Bereznyagovski forest
Russia (European) Voroninski Nature Reserve
Russia (European) Ichalkovski
Russia (European) Kulikovski forest
Russia (European) Tsninski Forest
Russia (European) Mouth of Svir river
Russia (European) Upper Voronezh forest
Russia (European) Neman river Delta and the coast of the Curonian Bay
Russia (European) Kraka Mountain
Russia (European) Central Meschera lake system
Russia (European) Flood-plain of Iput' river in vicinity of Kholevichami
Russia (European) Desna flood-plain near Trubchevsk
Russia (European) Okski Biosphere Reserve
Russia (European) Valley complex of Moksha and Oka rivers
Russia (European) Shilovo flood-plain of Oka river
Russia (European) Izhevsk flood-plain of Oka river
Russia (European) Solotcha flood-plain of Oka river
Russia (European) Watershed of Tsna and Vysha rivers
Russia (European) Moksha valley in vicinity of Temnikov
Russia (European) Moksha flood-plain in vicinity of Krasnoslobodsk
Russia (European) Flood-plain of Alatyr' river in vicinity of Ardatov
Russia (European) Kaluzhskiye Zaseki Nature Reserve
Russia (European) Nizhnekamskaya flood-plain
Russia (European) Watershed of Bel'skaya and Nugush rivers
Russia (European) Kamsko-Yayvenski wetland
Russia (European) Bel'skaya flood-plain
Russia (European) Zhigulevski Nature Reserve
Russia (European) Yamantau mountain
Russia (European) Kumikushski wetland
Russia (European) Iremel'ski mountain
Russia (European) Maly Nakas ridge
Russia (European) Irendyk ridge
Russia (European) Rakovye lakes
Russia (European) Flood-plain of Algashka river (Kirsko-Algashinskaya)
Russia (European) Dedinovo flood-plain of Oka river
Russia (European) Imeretinskaya lowland
Russia (European) Watershed between Sura and Barysh rivers
Russia (European) Kenozer'ye
Ukraine Kyivs'ke reservoir
Ukraine Bondar marsh
Ukraine Forests in Dnipro and Desna valleys
Ukraine Polis'kyi Nature Reserve
Ukraine Carpathian Biosphere Reserve
Ukraine Bodakivs'ke marsh
Ukraine Chervone marsh
Ukraine Udaj river valley
Ukraine Kanivs'kyi Nature Reserve
Ukraine Dniprovs'ko-Oril's'kyj Nature Reserve
Ukraine Syra Pogonya mire
Ukraine Pryp'yat' river valley
Ukraine Petropavlivs'kyj lyman
Ukraine Shats'ki lakes
Ukraine Turiya river valley
Ukraine Stokhod river valley
Ukraine Korotchenkivs'ki meadows
Ukraine Lyman lake
Ukraine Bagerove
Ukraine Ingul river valley
Ukraine Samars'kyj forest
Ukraine Mikulinetskie fish-ponds
Ukraine Oril' river valley
Ukraine Mzha river valley
Ukraine Chonovyny marsh
Ukraine Latorytsya river valley near Chop
Ukraine Psel river valley near Komsomol's'k
Ukraine Izyums'ka Luka forest
Ukraine Bakhtyn river valley
Ukraine Dereluj river valley
Ukraine Korostushevskoe Game Reserve
Ukraine Radomyshl's'ke Game Reserve
Ukraine Pischans'ke and Mogryts'ke forests
Ukraine Samara river valley
Ukraine Zakhidnyj Bug river valley
Ukraine Novograd-Volunskoe Game Reserve
Ukraine Ushomyrs'ke Game Reserve
Ukraine Solonyj Lyman lake
Ukraine Kremenets'kyj forest
Ukraine Orikhivski lake-system
Ukraine Psel river valley (Sary village)
Ukraine Meadows near Prociv village
Ukraine Debal'tsivs'ke lake
France Vallée de la Loire : gorges de la Loire
France Les Couzes Nord
France Baie des Veys et Marais du Cotentin
France Etangs d'Argonne
France Lac du Der-Chantecoq et étangs latéraux
France Vallée de l'Aube, de la Superbe et Marigny
France Vallée de l'Aisne
France Confluent des vallées de la Meuse et de la Chiers
France Bassigny
France Vallée de l'Yèvre
France Vallée de la Loire : confluence Loire-Vienne
France Basse vallée de L'Indre
France Vallée de la Loire : environs de Montlouis-sur-Loire
France Vallée du Fouzon
France Bassin du Drugeon: Pontarlier-Frasne
France Vallée de la Saône de Corre à Broye
France Basse Vallée du Doubs : Dole Sud
France Estuaire et embouchure de la Seine
France Boucle de Moisson
France Vallée de la Meuse
France Parc National des Ecrins
France Vallée de la Charente et de la Seugne (Cabariot-Pons/St-Sever-de-Saintonge)
France Vallée de la Charente : amont d'Angoulême
France Vallée de l'Oise de Thourotte à Vendeuil
France Estuaire de la Loire
France Basses Vallées Angevines
France Vallée de la Loire : de Nantes à Montsoreau
France Marais poitevin et baie de l'Aiguillon
France Val de Saône
France Parc National de la Vanoise
France Barthes de l'Adour
France Gorges du Gardon
France Etangs de Vendres, Pissevache et Lespignan
France Marais de L'Ile Vieille
France Marais de Brière
France Lac de Grand-Lieu
United Kingdom Lower Derwent Valley
United Kingdom Eden Estuary, Tentsmuir Point and Abertay Sands
United Kingdom Laggan Peninsula
United Kingdom Lewis Peatlands
United Kingdom Gruinart Flats
United Kingdom Mill Dam, Shapinsay
United Kingdom North Colonsay and Western Cliffs
United Kingdom North Uist Machair and Islands
United Kingdom Rum
United Kingdom Rinns of Islay
United Kingdom Uists Machairs, Lochs and Coast
United Kingdom South-eastern Stronsay
United Kingdom Tiree and Coll
United Kingdom Treshnish Isles
United Kingdom Ornkey Mainland Moors
United Kingdom Eoligarry, Barra
United Kingdom West Coast, Lewis (Corncrake)
United Kingdom Canna and Sanday
United Kingdom Cuillins
United Kingdom Gigha Island and Islets
United Kingdom South Walls and Switha
United Kingdom Stornoway to Back, Lewis
United Kingdom West Coast of Benbecula
Italy Tarvisio Forest
Italy Piave river
Italy River Biferno
Italy Assa valley
Italy Mount Grappa
Italy Belluno Dolomites
Italy Carniche pre-Alps
Italy Cividalese and upper Torre valley
Italy Susa and Chisone valleys
Italy Middle valley of Tagliamento river
Italy Giulie Pre-Alps
Luxembourg Aspelt - Lannebur, am Kessel
Luxembourg Upper Valley of the Alzette
Czechia Krkonose mountains (Giant mountains)
Czechia Confluence of the Morava (March) and Dyje (Thaya) rivers - Tvrdonicko
Czechia Poodri
Czechia Beskydy mountains
Czechia Jeseniky mountains
Czechia Doupov hills
Romania Defileul Inferior al Mureşului
Denmark Sejerø Bay and Nekselø
Denmark Lillebælt
Denmark Tystrup-Bavelse Søerne
Denmark Holmegård Mose & Porsmose
Denmark Ho Bugt meadows
Denmark Ribe Holme and meadows at Kongeåen
Denmark Tøndermarsken, Magisterkog and Rudbøl Sø
Denmark Sydfynske Ø-hav
Denmark Western part of Vejlerne, Arup Holm and Hovsør Røn
Denmark Hjarbæk Fjord
Denmark Almindingen
Denmark Bøtø Nor
Denmark Ballum og Husum Enge, Kamper strandenge
Denmark Frøslev Plantage and Frøslev Mose
Denmark Store Vildmose, Ryå and Stavad Enge
Georgia Eastern Caucasus
Georgia Iori Region
Georgia Javakheti Plateau
Georgia Kolkheti
Serbia Srednje Potamišje
Serbia Tara mountain
Serbia Uvac-Milesevka
Serbia Kopaonik
Serbia Prokletije
Slovakia Nízke Tatry
Slovakia Tatry
Slovakia Slovenský karst
Slovakia Slanské hills
Slovakia Horná Orava
Slovakia Senné
Slovakia Volovské hills
Slovakia Morava
Slovakia Riečnica
Slovakia Malá Fatra
Slovakia Chočské hills
Slovakia Bukovské hills
Slovakia Veľká Fatra
Slovakia Poiplie
Slovakia Poľana
Slovakia Medzibodrožie
Slovakia Lešť
Slovakia Levočské hills
Slovakia Spišská Magura
Slovakia Muránska plateau and Stolické hills
Slovakia Vihorlat
Slovakia Malé Karpaty
Serbia Sara mountain
Serbia Derdap gorge
Norway Nordre Øyeren & Sørumsneset
Lithuania Valley of Minija River
Belarus Bielaviežškaja Pušča
Denmark Marstal Bugt and the coast of south-west Langeland
Austria March/Thaya riverine forest
Austria Vienna forest
Austria Southern Waldviertel
Austria Allentsteig military training area
Austria Fish-ponds in the Waldviertel
Austria Western Waldviertel
Austria Niederösterreichische Randalpen
Austria Machland
Austria Freiwald
Austria Bohemian forest and Mühl valley
Austria Salzach valley
Austria Styrian Joglland
Austria Styrian Enns valley
Austria Meadows on the lower Ill
Austria Austrian part of Hanság
Austria Feuchte Ebene and Rauchwarther Platte
Slovakia Sysľovské fields
Slovakia Považský Inovec
Slovakia Tríbeč
Slovakia Danube floodplains
Poland Vistula river mouth
Switzerland La Brévine and Les Ponts-de-Martel
Switzerland Lower Engadine and Swiss National Park
Germany Erzgebirgskamm near Satzung
Germany Lower Oder valley
Germany Lower Havel - Lake Schollene - Lake Gülpe
Germany Middle Elbe valley with Steckby-Lödderitz forest
Germany Peenetal (Peenetalmoor and Anklamer Stadtbruch)
Germany Chiemsee and Chiemseemoore
Germany Danube valley: Regensburg-Vilshofen
Germany Danube valley: Neu-Ulm - Lauingen incl. Faiminger Stausee, Donau-Moos, and Gundelfinger Moos
Germany Rötelsee-Weihergebiet incl. Regenaue and Chambaue
Germany Federseeried
Germany Barnbruch
Germany Steinhuder Meer
Germany Pinneberg Elbe lowlands
Germany Aland-Elbe lowlands
Germany Drömling (Sachsen-Anhalt)
Germany Milde lowlands/Altmark
Germany Wulfener Bruch and Teichgebiet Osternienburg
Germany Elbe lowlands Jerichow
Germany Saale-Elster valley
Germany Rhön Biosphere Reserve
Germany Hellwegbörde
Germany Drömling (Niedersachsen)
Germany Helme reservoir Berga-Kelbra
Germany Main valley: Eltmann - Hassfurt
Germany Altmühl valley between Treuchtlingen and Leutershausen with Altmühlsee
Germany Spreewald
Germany Donau-Auen and Donau-Ried: Höchstädt - Donauwörth
Germany Döberitzer Heide
Germany Oberes Rhinluch / Havelländisches Luch Nauen-Friesack
Germany Eschefeld ponds
Germany Eastern Erzgebirge around Fürstenau
Germany Wagbach lowlands
Germany Borgfelder Wümmewiesen
Germany Blockland - lower Wümme valley - Westliches Hollerland
Germany Werderland
Germany Niedervieland, Ochtumniederung
Germany Rietzer See
Germany Peitzer und Bärenbrücker Teiche
Germany Uckermark lake region
Germany Lower Elbe valley
Germany Deichvorland Oderbruch
Germany Unteres Rhinluch-Dreetzer See / Havelländisches Luch / Belziger Landschaftswiesen
Germany Schorfheide-Chorin
Germany Lower Rhine
Germany Heubachniederung/ Schwarzes Venn/ Borkenberge/ Halterner Seen
Germany Wet grasslands and forests around Burbach
Germany Medebacher Bucht
Germany Lower reaches of River Weser, unembanked area
Germany Winsener Elbmarsch
Germany Leine valley near Salzderhelden
Germany Valley of River Ems
Germany Südradde lowlands
Germany Old Picardie
Germany Lower Aller lowlands
Germany Ostenholzer Moor and Meissendorfer Teiche
Germany Dümmer
Germany Hamme lowlands
Germany Elbe lowlands between Schnackenburg and Lauenburg
Germany Elbe marshes between Stade and Otterndorf
Germany Wümme lowlands and St. Jürgensland
Germany Wümmewiesen near Fischerhude
Germany Wendesser moor
Germany Heerter See
Germany Lowlands of middle Havel
Germany Aisch-Regnitz-Grund
Germany Murnauer Moos and Loisach-Kochel-Moore
Germany Fiener Bruch
Germany Rheiderland
Germany Ems valley between Leer and Emden
Germany Lowlands of the Rivers Eider, Treene and Sorge
Germany Hudewälder northeast of Haldensleben
Germany Duvenstedter Brook
Germany Lahn valley from Marburg to Wetzlar
Germany Bliesgau
Germany Alluvion of the middle Horloff
Germany Hessian Rhine alluvion
Germany South-eastern Alb and Upper Danube valley
Germany Enz valley Mühlhausen - Roßwag
Germany Woodlands of Western Pomerania, northern range
Germany Feldberg-Woldegk terminal moraine
Germany Heidmoor lowlands
Norway Jæren
Luxembourg Valley of the Ernze Blanche
Croatia Vrana Lake and Jasen
Croatia Poilovlje with Carp Fishponds
Croatia Lower Posavina
Croatia Turopolje
Jordan Northern Jordan Valley (North Ghor)
Croatia Alluvial wetlands of the River Danube
Germany Recker Moor/ Düsterdieker Niederung/ Seester Feld
Montenegro Durmitor
Botswana Okavango Delta
Egypt Lake Bardawil
Egypt Zaranik Protected Area
Egypt Lake Manzala
Kenya Tsavo West National Park
Kenya Masai Mara
Tanzania Ugalla River Game Reserve
South Africa Kruger National Park and adjacent areas
South Africa Grasslands
South Africa Mkuzi Game Reserve
South Africa Chelmsford Nature Reserve
South Africa Hluhluwe-iMfolozi Park
South Africa Umlalazi Nature Reserve
South Africa Maloti Drakensberg Park
South Africa Umvoti Vlei
South Africa KwaZulu-Natal Mistbelt Grasslands
South Africa Franklin Vlei
Zambia Hillwood
Zambia Nkanga River Conservation Area
Zambia Kafue Flats
Zambia Kafue National Park
Zambia Chisamba
Zambia Nyika National Park (Zambia)
Zambia Shiwa Ng'andu
Estonia Räpina polder
Latvia Ziemelgauja
Lithuania Flooded coastal meadows near Tyrai bog
Lithuania Zemaitija National Park
Lithuania Flooded meadows arround the Senoji & Naujoji Rusne lakes
Ukraine Tsybul'kivs'ki lakes
Ukraine Karachunivs'ke reservoir
Ukraine Mishuryn Rig
Ukraine Uzh river valley
Ukraine Gajchur river valley
Lithuania Birveta Fish-farm including segments of Dysna River and Birveta River valleys
Belarus Sporaŭskaje balota
Ukraine Yavoriv Military Training Centre
North Macedonia Lake Dojran
Netherlands IJssel: IJsselkop - Ketelmeer
Poland Pasleka river valley
Ukraine Styr' river valley (Kolky village)
Ukraine Velyka Osokorovka
Ukraine Berezhnytsya river valley
Ukraine Male Polissya
Ukraine Vyshnja river valley
Ukraine Crimean Nature Reserve
Ukraine Velika babka river valley
Ukraine Snake island
Ukraine Dniprodzerzhyns'ke reservoir
Ukraine Sejm river valley (Mutyn village)
Ukraine Sejm river valley (Vyrky village)
Ukraine Gomol'shans'ka forest
Russia (European) Vicinity of Dakhovka
Russia (European) Kaltasinski forest
Russia (European) Krasnokamski forest
Russia (European) Aiskiye yary
Russia (European) Mountain valley of Zilim river
Russia (European) Sondugski Reserve and its outskirts
Russia (European) Druzhinnoe lake and its outskirts
Russia (European) Vicinity of the Vozhe lake and Charonda marshes
Russia (European) Fish ponds in the vicinity of Kovashi and Syur'yevskoye bog
Russia (European) Klyazminski reserve
Russia (European) Flood-plain of Lukh river between Myt and Mugreevo-Nikol'skoye villages
Russia (European) Valley of Zhizdra river
Russia (European) Bryansko-Zhizdrinskoye woodland
Russia (European) Watershead of Atsvezh and Yuma rivers
Russia (European) Flood-plain of Vad river
Russia (European) Flood-plain of Sura river
Russia (European) Homeland of the Crane (Dubna marshes and adjacent areas)
Russia (European) Siz'menski flood-plain of Sheksna reservoir
Russia (European) Polesski Forest
Russia (Asian) Belozersk lakes
Russia (Asian) Tunkin valley
Belarus Balota Dzikoje
Belarus Prostyr
Belarus Biarezina-Hajna
Belarus Brody
Belarus Servač
Belarus Liebiadziny Moch
Belarus Cikiny astravy
Belarus Dražbitka-Svina
Belarus Janka river floodplain
Belarus Prypiackija baloty
Estonia Haanja
Estonia Karula
Estonia Võrtsjärv
Estonia Lahemaa
Estonia Väinameri
Georgia Alazani Valley
Georgia Lower Kura Valley
Georgia Kartsakhi Lake
Georgia Tabatskuri Lake
Georgia Khanchali Lake
Georgia Madatapha Lake
Georgia Pharavani Lake
Georgia Sagamo Lake
Georgia Bogdasheni Lake
Georgia Kazbegi
Russia (European) Sviyago-Kubninskaya forest-steppe
Russia (Central Asian) Tobolo-Kurtamyshskay flood land
Russia (European) Sokolino-Kasplyanskaya area
Russia (European) Samarskaya Luka
Russia (Central Asian) Bystroistokskaya
Russia (Central Asian) Bolchoe Kamyshnoe Lake
Russia (Central Asian) Zonalnay
Russia (Central Asian) Mouth of the Uy river
Russia (Central Asian) Middle reaches of the Chulym river
Russia (Central Asian) Bol'shoye Beloye lake
Russia (Central Asian) Kisluhinskaya
Russia (Central Asian) Busly lake
Russia (Central Asian) Teletskoye lake
Russia (Central Asian) Pelymsky Tuman
Russia (Central Asian) Visimsky zapovednik and vicinity
Russia (Central Asian) Zapovednik "Denezhkin Kamen'"
Russia (Central Asian) Chernoye Lake
Russia (Central Asian) Oleni brooks
Russia (Central Asian) Dikoye and Epanchino lakes
Russia (Central Asian) The top part of pools of the rivers Big Karaganka and the rivers Syntasty
Russia (Central Asian) Flood-plain of Tobol river near Lebedevka and Bugrovoye villages
Russia (Central Asian) Natural reserve Kuznetskij Alatau
Russia (Central Asian) Flood-plain of Tobol river between mouths of the Uya and Ubagan rivers
Russia (Central Asian) Yel'tsovskaya
Russia (Central Asian) Shestakovskie of a bog 2
Russia (Central Asian) Ilmensky zapovednik
Russia (Central Asian) Channel Siberian
Russia (Central Asian) Kileinoye bog
Russia (Central Asian) Vicinities of lake Achikul
Czechia Labské pískovce sandstones
Czechia Orlicke Zahori
Czechia Kralicky Sneznik
Czechia Libava
Russia (Central Asian) Charyshskaya
Russia (Central Asian) Krasnoschekovskaya
Russia (Central Asian) Valley of the Berd' river
Russia (Central Asian) Anuyskaya
Russia (Central Asian) Vaganovskaya
Russia (Central Asian) Yel'tsovskaya
Russia (Central Asian) Watershed of the Mulym'ya and Bolshoy Tap rivers
Russia (Central Asian) Taezhno-Mikhaylovsky
Russia (Central Asian) Kurtan Lake
Russia (Central Asian) Siberian Channel
Russia (Central Asian) Flood-plain of the Tuy river
Russia (Central Asian) Ust'-Ozerninskiye bogs
Russia (Central Asian) South spurs of Sylvinski range
Russia (Central Asian) Sources of the Bolshaya Karaganka and Syntasty rivers
Russia (Central Asian) Turina mountain
Russia (European) Karakulinskaya flood-plain
Russia (Central Asian) Cheka mountain
Russia (Central Asian) Kazanskaya flood-plain of the Ishim river
Russia (Central Asian) Marsh westward of village Stepanovka near Okunevo Lake
Russia (Central Asian) Kataiginskiye bogs
Russia (Central Asian) Kuznetsky Alatau Zapovednik
Russia (Central Asian) Flood-plain of river Irtysh near village Atak
Russia (Central Asian) Shestakovskiye marshes
Latvia Abava valley
Latvia Abeli
Latvia Dubna flood-plain
Latvia Garkalne forest
Latvia Gauja national park
Latvia Teici and Pelecare bogs
Latvia Adazi
Latvia Seda marsh
Latvia Irbe valley and Dizpurvs
Latvia Augsdaugava
Latvia Kuja
Latvia Kemeri national park
Latvia Pededze forest and Parabaine
Latvia Vecumi
Latvia Zvarde forest
Latvia Stikli
Latvia Sventaja valley
Latvia Babite lake
Latvia Piejura
Latvia Rakupe and Pace
Latvia Burga meadows
Latvia Burtnieks' meadows
Latvia Mezole
Latvia Randu meadows
Latvia Sturu and Zvardes bogs
Latvia Razna
Latvia Tasu lake
Latvia Salaca valley
Georgia Adjara-Imereti Ridge
Bulgaria Ivailovgrad Reservoir
Bulgaria Rupite
Bulgaria Kresna Gorge
Bulgaria Kaliakra
Bulgaria Sakar
Bulgaria Emine
Bulgaria Batova
Bulgaria Sredna Gora
Bulgaria Rila
Bulgaria Pirin
Bulgaria Ludogorie
Bulgaria Western Strandzha
Bulgaria Karlukovski Karst
Bulgaria Dobrostan
Bulgaria Osam Meadows
Bulgaria Kamchiyska Mountain
Russia (Central Asian) Sibirskaya anabranch (Irtysh flood-plain)
Armenia Haghartsin
Armenia Jermook
Armenia Mount Ara
France Bassin d'Arcachon et Banc d'Arguin
France Anse de Fouras, baie d'Yves et marais de Rochefort
France Anse du Fiers d'Ars en Ré
France Iles du Haut-Rhône
France Ried de Colmar et Sélestat
France Planèze de Saint-Flour
France Brenne: Forêt de la Petite-Brenne
France Bassée et plaines adjacentes
France Plateau de Delme, Val de Petite Seille
France Estuaire de la Charente
France Plaine calcaire du sud Vendée
Bulgaria Nikopolsko Plateau
Kazakhstan Big Almaty Gorge
Kazakhstan Aksu-Dzhabagly State Nature Reserve
Russia (European) Flood-plain of Oka river in Vachski district of North Novgorod region and adjacent areas of Vladimir region
Russia (European) Ichalkovski forest
Russia (European) Shumerlinskaya
Russia (European) Floodplain of Pjana river near village Sarga
Russia (European) Floodplain of Moksha river and Common Crane autumn aggegation near villages Khohlikha and Shaprikha
Russia (European) Flood-plain of river Vetluga
Russia (European) Pizhemski reserve
Russia (European) Forests and bogs in the lower reaches of Bol'shaya Kaksha river
Russia (European) Nothern part of Gor'koe reservoir with spurs
Russia (European) Flood-plain of rivers Usta and Vaya in Urenski district
Russia (European) Kilemarski Reserve
Russia (European) Kurmyshskaya flood-land
Russia (European) Forest in the middle reaches of Serezha river
Russia (European) Semilovskoye forestry
Russia (European) Dono-Donetskoye flood-plain
Russia (European) Floodplain of Rudnya river and surroundings
Russia (Central Asian) Ata-Anay Lake
Russia (Central Asian) Seketovo, Rakhtovo and Artevo lakes
Russia (Central Asian) Pershinsko-Manatkinsky area
Russia (Central Asian) Baturino-Simansky area
Russia (Central Asian) Ubinskoye Lake
Russia (Central Asian) Zaikovo forest
Russia (Central Asian) Mayan lake
Russia (Central Asian) Bol'shoy and Maly Vagilsky Tuman
Iran, Islamic Republic of Alborz-e Markazi Protected Area
Iran, Islamic Republic of Lashgarak and Latian dam
Iran, Islamic Republic of Golestan
Lithuania Wet meadows of Namunas River valley near Viesvile
Israel Hula valley
Israel Zevulun valley
Israel Carmel coast
Israel Jezre'el, Harod and Bet She'an valleys
Israel Hefer valley
Israel Judean foothills
Israel Judean desert
Israel Western Negev
Israel Northern Arava valley
Israel Southern Arava valley and Elat mountains
Jordan Jordan Valley
Jordan Madaba - Hisban
Kuwait Al-Jahra Pool Nature Reserve
Lebanon Aammiq wetland
Lithuania Valley of Nemunelis River
Lithuania Valley of Venta River
Czechia Boletice
Oman Masirah island
Palestine Jerusalem wilderness
Saudi Arabia Gulf coral islands
Lithuania Kamanos Strict Nature Reserve
Saudi Arabia Al-Ha'ir
Saudi Arabia Mahazat as-Sayd
Saudi Arabia National Wildlife Research Center and environs, Taif
Yemen High mountains of Ibb
Bulgaria Vasilyovska Mountain
Bulgaria Ponor
Bulgaria Kotlenska Mountain
Bulgaria Palakaria
Bulgaria Ovcharovo Meadows
Bulgaria Ruy Mountain
Bulgaria Boboshevo Meadows
Bulgaria Noevtsi Meadows
Bulgaria Vitosha Mountain
Bulgaria Dolna Koznitsa Meadows
Bulgaria Rayanovtsi
Bulgaria Central Balkan
Bulgaria Mikre Meadows
Bulgaria Apriltsi Meadows
Bulgaria Berkovitsa Meadows
Bulgaria Kocherinovo Meadows
Bulgaria Velchevo Meadows
Bulgaria Meshtitsa Meadows
Bulgaria Skrino
Bulgaria Dolni Bogrov-Kazichene
Bulgaria Straldzha Complex
Bulgaria Devetashko Plateau
Uzbekistan Aksay Lake and surrounding desert
Austria North-eastern Leithagebirge
Austria Surroundings of Mattersburg
Italy Valle del Torrente But
Italy Tarvisio forest and Giulie pre-alps
Austria Parndorfer Platte and Heideboden
Austria Zoological Gardens of Lainz
Austria Riverine forests on the Danube east of Vienna
Austria Middle Kamp valley
Austria Steinfeld
Austria Thermenlinie
Austria Meadows and lakes in the foreland of the Alps in Salzburg and Upper Austria
Austria Ibmer Moor
Austria Northern Kalkalpen
Austria Niedere Tauern
Italy Carniche Alps
Italy Lagorai mountains
Austria Lower valley of the Gail
Ukraine River Tysa Valley
Ukraine Chornobyl zone
Ukraine Urochysche Mushevo
Ukraine Seredinno-Budskyj Park
Ukraine Desna river part of National Nature Park "Desnyans'ko-Staroguts'kyj"
Ukraine Regional Landscape Park Trakhtemyriv
Ukraine Serednyotomakivske reservior
Ukraine Ivanivka vallage
Ukraine Ai-Petri
Austria Riede in the northern Rhein valley
Ukraine National Park Stara Guta
Ukraine Regional Landscape Park Mal'ovanka
Ukraine Vodorazdil'ni i Zovnishni Gorgany
Ukraine Teteriv river valley
Ukraine Buzhok river valley (Anastavtsi)
Ukraine Reservoir on r. Chechva with territories adjacent
Ukraine Upper reaches of r. Sivka
Ukraine Vorskla river valley, near vv. Ivanivka-Kashans'ke, Kurikovka
Ukraine Komisarovskyj Forest
United Kingdom Tiree (Corncrake)
United Kingdom Coll (Corncrake)
United Kingdom Iona
Ukraine Upper of r.Mokra Sura
Kazakhstan Western and northern foothills of the Kalba Range
Ukraine Sinelnikov Complex
Sweden Lake Lim
Poland Barycz river valley
Kazakhstan Almaty State Nature Reserve
Poland Gorce mountains
Poland Weltyn lakes
Poland Wkra and Mlawka Rivers Valleys
Russia (European) Bolchoi Tkhach mountain
Russia (European) Sources of Kuna and Shisha rivers
Russia (European) Sources of Kuma river
Russia (European) Ravine of Eshkakon river
Russia (European) Sources of the Podkumok river
Russia (European) Marinskaya cuesta of Skalisti ridge
Sweden Sjöängen – Svartån
Russia (European) Digoriya rocks
Russia (European) Malka river ravine
Russia (European) Ravine of Gundelen-Tyzyl river
Russia (European) Sources of Khudes river
Poland Wkrzanska Forest
Poland Cedynia Site
Poland Goleniow Forest
Poland Trzebiatow Site
Poland Witnica-Debno Site
Russia (European) Akhmet-Skala ridge
Poland Barlinek Forest
Poland Drawa River Site
Poland Forest at Drawa River
Poland Middle Odra River Valley
Poland Rogalin Site
Poland Goplo Lake Site
Poland Calowanie Swamp
Poland Bielawa Swamps
Poland Darzlubska Forest
Poland Mirachowo Forest
Poland Lower Silesia Forests
Poland Pilica River Valley
Poland Kozienicka Forest
Poland Kostrzyn River Valley
Poland Warmia Site
Poland Dobskie Lake
Lebanon Hima Ebel es-Saqi
Poland Skaliskie Forest
Poland Orzysz Military Area
Poland Wizna Swamps
Poland Upper Nurzec River Valley
Poland Lower Notec River Valley
Poland Middle Notec River Valley
Poland Notec Forest
Poland Podedworze Reservoir
Poland Mosty-Zahajki Site
Poland Polesie
Poland Por River Valley
Poland Nielisz Site
Poland Tyszowce Site
Poland Upper Huczwa River Catchment
Italy Orobie Alps and pre-Alps
Poland Solokija River Valley
Italy Grigne
Poland Stolowe Mountains
Poland Upper Labunka River Valley
Poland Lower Skawa River Valley
Poland Orawa and Nowy Targ Bogs
Poland Beskid Niski Mountains
Poland Slonne Mountains
Kazakhstan Paradise Valley mountain plateau
Russia (European) Gunibskoye plateau
Lebanon Cheikh Zennad
Lebanon Horsh Ehden Nature Reserve
Lebanon Palm Islands Nature Reserve
Poland Lower Narew River Valley
Netherlands Lake Sneekermeer and Goingarijp
Lebanon Shouf Cedars Nature Reserve
Romania Avrig - Scorei - Făgăraş
Slovenia Breginjski Stol and Planja
Slovenia Kozjansko - Jovsi
Russia (Central Asian) Upper streams of Ob' river
Slovenia Snežnik plateau and Pivka valley
Slovenia Banjšice plateau
Slovenia Dravinja valley
Poland Narew river gaps
Slovenia Vipavski rob
Russia (Asian) Sayanski canyon of the Enisey river
Russia (European) Sochinsky National Park
Russia (Asian) Islands of the Enisey river near Shushenskoye
Russia (Asian) Perovo lake
Netherlands De Wieden
Poland Solska Forest
Poland Bzura River Valley
United Kingdom Nene Washes
Hungary Borsodi-Mezőség
Hungary Alkaline pusztas of Upper Kiskunság
Hungary Upper Tisza
Slovakia Laborecká upland
Armenia Pombak mountain chain
Croatia Lika Karst Fields
Croatia Plitvice Lakes National Park
Croatia Cetina River
Ireland North west Connemara (Cleggan, Inishbofin, Inishturbot, Omey)
Ireland Inishsirrer, Inishmeane, Gola & Owey Islands
Finland Oulu region wetlands
Finland Outokumpu wetlands
Finland Päätyeenlahti bay
Finland Laajalahti bay, Vanhankaupunginlahti bay and Viikki
Russia (Central Asian) Schuch'ye lake
Belarus Arechaŭskaje
Belarus Dniepr floodplain Lojeŭ-Žary
Belarus Čyrvony Bor
Germany Schleswig-Holstein Wadden Sea National Park (includes the Halligs, Kniepsand/Amrum, Lister Koog, Rantumbecken, Rickelsb
Poland Upper Vistula River Valley
North Macedonia River Radika catchment
Germany Forests of Lauenburg
Russia (European) Pikhtovka fishponds
Russia (European) Floodplain of Cheptsa river
Russia (European) Tul'skiye Zaseki forest
Russia (European) Zalazniki, Beli, Urino and Leskovo area
Russia (European) Floodplain of Ural and Kindeli rivers
Russia (European) Rychkovskaya forest-steppe
Russia (European) Watershed of Kazanka and Shoshma rivers
Russia (European) Karabash-Kudashskaya forest-steppe
Russia (European) Pis'myanskiye Gori area
Russia (European) Downstream of Ik river
Russia (European) Kundryuchenskiye sands
Russia (European) Severo-donetskaya
Russia (European) Rasskazan' lake and flood-plain of Khoper river
Russia (European) Prisyun'ski forest
Russia (European) Buninski forest
Russia (European) Manturovo left bank of the Unzha river
Russia (European) Floodplains of the Unzha river near Kologriv
Russia (European) Kologrivski forest
Russia (European) Vetluga floodplain in the Nizhni Novgorod region
Russia (European) Tishskaya Dal' area
Germany Upper and middle reaches of river Krückau with tributaries
Russia (European) Spassk flood-plain of Oka river
Russia (European) Forests and wetlands in the valley of the Nuzhma river
Russia (European) Klyazminsko-Balakhninskaya lowland
Russia (European) Vedeneevsk pine forest
Russia (European) Flood-plain of Klyaz'ma river from mouth of Teza river up to Mstera
Russia (European) Flood-plain of Lukh river between Lukh and Myt villages
Germany Hunte lowlands near Oldenburg
Germany Leeg-, Melm- and Kuhdammoor
Germany Elzniederung between Kenzingen and Rust
Germany Ostfriesische Meere
Germany Fehntjer Tief
Germany Isar valley: Gottfrieding - Plattling incl. Isar mouth
Germany Ammersee
Germany Lakes of Schwerin, Dambeck and Warin
Germany Rodachaue / Itzgrund / Oberes Maintal incl. Nassanger near Trieb and surrounding gravel pits
Germany Viehmoor with Leiferder Teiche
Germany Alfsee
Germany Stepenitz-Maurine-Radegast
Germany Upper and middle reaches of river Warnow with Göwe and Mildenitz
Germany Lower reaches of river Warnow
Germany Mecklenburgische Schweiz
Germany Recknitz and Trebel valley
Germany Einswarder Plate / Tegeler Plate
Germany Geeste lowlands near Bramel
Germany Huvenhoopsmor and Breddorfer Wiesen
Germany Drennhauser Hinterdeich
Germany Mittleres Wietingsmoor
Germany Georgsdorfer and Dalum-Wietmarscher Moor and Alte Piccardie
Germany Oderbruch
Germany Oder lowlands Lebus-Ratzdorf
Germany Randow-Welse-Bruch / Uckermärkische Agrarlandschaft
Germany Mittlere Schwäbische Alb with Albuch, Härtsfeld and Schmiechener See
Germany Danube valley Scheer-Zwiefaltendorf
Germany Osterried near Hermaringen
Germany Wurzacher Ried
Germany Main valley and Steiger foothills near Schweinfurt and Volkach
Germany Spessart
Germany Island of Usedom
Germany Putzarer See, Galenbecker See, Brohmer Berge
Germany Ueckermünder Heide
Germany Elbe valley of Mecklenburg
Germany Schönbuch with Spitzberg
Germany Vorland of the Mittlere Schwäbische Alb
Germany Bodanrück and Mindelsee
Germany Untersee of Lake Constance
Germany Baar
Germany Donauried
Germany Vogelsberg
Germany Wetterau
Germany Schwalmaue around Schwalmstadt
Germany Werra lowlands near Wanfried and Eschwege
Germany Mönchbruch of Mörfelden and Rüsselsheim and heath extensions (GG)
Germany Lower Gersprenz lowlands incl. Reinheim pond
Germany Obersuhl lowlands
Germany Wümme lowlands Borgfeld-Fischerhude
Germany Lüneburger Heide
Germany Moorland belt of Süderelbe-Marschrandmoor (HH) and moorlands near Buxtehude (NI)
Germany Serrahn and Woldegk-Feldberger Hügelland
Germany Elbe lowlands near Bertingen
Germany Elbe meadows at Großes Ostragehege Dresden
Germany Wurzelbergfarmde
Germany Nuthe - Nieplitz lowlands
Germany Lauenburgische Seen Nature Park and Schaalsee area
Germany Bayerischer Wald National Park with Arber region and Hohem Bogen
Germany Werra meadows near Bad Salzungen
Germany Bastau lowlands
Germany Salziger See and Salza valley
Germany Mouth of river Schwarze Elster
Germany Ahrmündung
Germany Engerser Feld
Germany Mittelrheinisches Becken with Laacher See and Thürer Wiesen
Germany Speyerer Wald, Nonnenwald and Bachauen between Geinsheim and Hanhofen
Germany Offenbacher Wald, Bellheimer Wald and Queichwiesen
Germany Bienwald and Viehstrichwiesen
Germany Kühren Teich and Lanker See
Germany Flensburger Innen- and Aussenförde
Germany Haaler Au lowlands
Ireland Inishsirrer, Inishmeane, Gola and Owey Islands
Poland Pulwy Marshes
Romania Câmpia Cermeiului
Romania Câmpia Crişurilor
Romania Câmpia Nirului - Valea lerului
Romania Nerei gorge - Beuşniţa
Romania Confluenţa Jiu - Dunăre
Romania Cursul Mijlociu al Someşului
Romania Dealurile Homoroadelor
Romania Dealurile Nordice ale Munţilor Făgăraş
Romania Dealurile Târnavelor - Valea Nirajului
Romania Ciuc Depression
Romania Giurgeu Depression
Romania Dumbrăviţa - Rotbav - Măgura Codlei
Romania Iazurile de pe Văile Ibănesei - Başeului - Podrigăi
Romania Lunca Turului Inferior
Romania Mures river lower flood-plain
Romania Timiş river flood-plain
Romania Mândra wetlands
Romania Baraolt Mountains - Bodoc
Romania Trascău Mountains
Romania Bârnova woodland
Romania Bogata forest
Romania Macedonia forest
Romania Piatra Şoimului - Scorţeni - Gârleni
Romania Podişul Hârtibaciului
Romania Fizes valley
Romania Râului Negru valley
Romania Vânători - Neamţ
Romania Balta Albă - Amara - Jirlău
Luxembourg Region of the Middle Lias
United Kingdom South Uist Machair and Lochs
Luxembourg Valley of the Syre
Kyrgyzstan Tokmak Pheasant Reserve
Poland Sieniawa Forest
Poland Beskid Wyspowy Mountains
Poland Czarna River Valley
Poland Lower San River Valley
Poland Lower Wieprz River Valley
Poland Wloszczowa Basin
Poland Mircze Site
Poland Poprad Site
Poland Sandomierz Forest
Poland Sudety Wałbrzysko-Kamiennogorskie Mountains
Poland Elblag Height
Hungary Alkaline lakes of Kiskunság
Denmark Jerup Hede, Råbjerg Mose and Tolshave Mose
Denmark Råbjerg Mile and surrounding heathlands
Denmark Sønder Ådal
Denmark Eastern part of Vejlerne
Lithuania Dysnai & Dysnykstis lake with adjacent territory
Lithuania Meadows of Nemunas River valley between Raudone & Gelgaudiskis
Lithuania Valley of Nevezis River
Hungary Zemplén Mountains
Lithuania Agricultural land near the Kalvarija settlement
Lithuania Dubysa River hereinafter Lyduvenai settlement & its valley
Lithuania Valleys of the Saltoja & Vyzuona rivers
Lithuania Segments of the Apse River valley
Lithuania Valleys of the Erla & Salantas rivers
Lithuania Svyla River valley
Poland Szczecin lagoon
Poland Kamien Lagoon and Dziwna River
Poland Tuchola Forest
Poland Knyszyn Forest
Poland Lukow Forests
Poland Mala Welna River Valley
Poland Pszczew Lakes and Obra River Valley
Poland Slawskie Lakes
Poland Woniesc reservoir
Kazakhstan Kultansor and Tatysor Lakes
Kazakhstan Tassuat Lake
Poland Strzeleckie forests
Poland Roztocze
Poland Szyszla River Valley
Poland Karkonosze Mountains
Poland Nysa reservoir
Poland Upper Odra River Valley
Poland Brzeszcze ponds
Poland Niepolomice forest
Poland Beskid Zywiecki Mountains
Poland Pieniny Mountains
Poland Beskid Slaski Mountains
Poland Grzeda Sokalska Beechwood
Poland Przysowia and Sludwia River valley
Poland Stobrawa Watershed
Poland Widawa and Olesnica River Valley
Poland Izerskie Mountains
Poland Radlow Forest
Poland Kurpie Site
Poland Kwiecewo Site
Poland Satopy-Samulewo Site
Poland Peatbogs at Wieprz River
Poland Jaktorow and Krasnicza Wola Fishponds
Poland Swietokrzyskie Vistula River Valley
Poland Irrigation Fields in Wroclaw
Poland Sredzka Struga and Maskawa River Valley
Poland Siemianowka Reseirvoir
Bosnia and Herzegovina Livno karst field and Busko lake
Slovenia Breginjski Stol
Slovenia Dobrava - Jovsi
Slovenia Krakovo forest and Šentjernej plain
Zambia Uningi Pans
Belarus Braslav lakes
Jordan Maghtas - Sweimeh
Armenia Amasia
Armenia Dsegh
Armenia Gndasar
Moldova Royal Forest
South Africa Penny Park
South Africa Suikerbosrand Nature Reserve
Serbia Vlasina
Serbia Pester
Russia (European) Yamskaya steppe and adjacent technical reservoirs
Denmark Bolle and Try meadows
Denmark Tinglev Mose and Sø
Estonia Karala-Pilguse
Estonia Kasti bay
Estonia Lõu bay
Estonia Kikepera
Estonia Koorunõmme
Estonia Irbe strait
Estonia Kõrgessaare-Mudaste coast
Estonia Kärevere
Estonia Mullutu
Estonia Nõva-Osmussaar
Estonia Pärnu bay (NEW)
Estonia Riksu coast
Estonia Siiksaare-Oessaare bays
Estonia Kunda
Estonia Vilsandi archipelago
Estonia Vooremaa
Croatia Gorski Kotar and Northern Lika
Croatia Pokuplje Basin
Estonia Agusalu
Greece Evros delta
Greece Kalloni Gulf, Lesvos
Greece Mount Olympos and pine forest of central Lesvos
Netherlands Rijntakken
Netherlands Uiterwaarden Zwarte Water & Vecht
Netherlands Drentsche Aa-gebied
Netherlands Oldambt
Estonia Kõnnumaa-Väätsa
Estonia Kõrvemaa
Estonia Käntu-Kastja
Estonia Küdema bay
Estonia Lavassaare
Estonia Marimetsa-Õmma
Estonia Meenikunno bog
Estonia Misso
Georgia Eastern Caucasus mountains
Armenia Dsegh - Haghartsin - Pombak chain and Dilijan National Park
Armenia Gorayk
Azerbaijan Barda tugai forest
Iran, Islamic Republic of Akh Gol
Iran, Islamic Republic of Aras Dam Lake
Iran, Islamic Republic of Lavandavil Wildlife Refuge
Iran, Islamic Republic of Anzali Mordab complex
Iran, Islamic Republic of Bojagh
Georgia Kvernaki Ridge
Kazakhstan Chokpak Pass
Kazakhstan Chingiztau Mountains
Kazakhstan Zhagalbayly and Tuyemoynak Hills
Kazakhstan Karabas Mountains
Serbia Gornji Visok and Vidlic
Serbia Ozren-Jadovnik
Serbia Svrljisko pobrdje
Netherlands Onlanden

Habitats & altitude
Habitat (level 1) Habitat (level 2) Importance Occurrence
Artificial/Terrestrial Pastureland suitable non-breeding
Artificial/Terrestrial Pastureland suitable breeding
Artificial/Terrestrial Plantations unset breeding
Grassland Subtropical/Tropical Seasonally Wet/Flooded major non-breeding
Grassland Temperate major breeding
Savanna Dry major non-breeding
Wetlands (inland) Bogs, Marshes, Swamps, Fens, Peatlands major breeding
Altitude 0 - 3000 m Occasional altitudinal limits  

Threats & impact
Threat (level 1) Threat (level 2) Impact and Stresses
Agriculture & aquaculture Annual & perennial non-timber crops - Agro-industry farming Timing Scope Severity Impact
Ongoing Majority (50-90%) Slow, Significant Declines Medium Impact: 6
Ecosystem degradation, Ecosystem conversion
Biological resource use Hunting & trapping terrestrial animals - Intentional use (species is the target) Timing Scope Severity Impact
Ongoing Minority (<50%) Negligible declines Low Impact: 4
Species mortality
Climate change & severe weather Storms & flooding Timing Scope Severity Impact
Ongoing Minority (<50%) Negligible declines Low Impact: 4
Ecosystem degradation, Reduced reproductive success
Human intrusions & disturbance Recreational activities Timing Scope Severity Impact
Ongoing Minority (<50%) Negligible declines Low Impact: 4
Species disturbance
Invasive and other problematic species, genes & diseases Invasive non-native/alien species/diseases - Felis catus Timing Scope Severity Impact
Ongoing Minority (<50%) Negligible declines Low Impact: 4
Reduced reproductive success
Invasive and other problematic species, genes & diseases Invasive non-native/alien species/diseases - Neovison vison Timing Scope Severity Impact
Ongoing Minority (<50%) Negligible declines Low Impact: 4
Reduced reproductive success
Invasive and other problematic species, genes & diseases Problematic native species/diseases Timing Scope Severity Impact
Ongoing Majority (50-90%) Negligible declines Low Impact: 5
Reduced reproductive success
Natural system modifications Other ecosystem modifications Timing Scope Severity Impact
Ongoing Majority (50-90%) Negligible declines Low Impact: 5
Ecosystem degradation
Transportation & service corridors Roads & railroads Timing Scope Severity Impact
Ongoing Minority (<50%) Negligible declines Low Impact: 4
Species disturbance

Recommended citation
BirdLife International (2022) Species factsheet: Crex crex. Downloaded from on 03/07/2022. Recommended citation for factsheets for more than one species: BirdLife International (2022) IUCN Red List for birds. Downloaded from on 03/07/2022.