Brandt's Cormorant Urile penicillatus


Acosta, S.; Jahncke, J.; Merkle, W.; Rachowicz, L. 2010. Ecological studies of seabirds on Alcatraz Island, 2010. Final report to the Golden Gate National Recreation Area National Park Service. PRBO Conservation Science, Petaluma, CA.

Butcher, G. S.; Niven, D. K. 2007. Combining data from the Christmas bird count and the breeding bird survey to determine the continental status and trends of North American birds.

Buxton R.T.; Galvan R.; McKenna M.F.; White C.L.; Seher, V. 2017. Visitor noise at a nesting colony alters the behavior of a coastal seabird. Marine Ecology Progress Series 570: 233-246.

del Hoyo, J.; Elliot, A.; Sargatal, J. 1992. Handbook of the Birds of the World, Vol. 1: Ostrich to Ducks. Lynx Edicions, Barcelona, Spain.

Delany, S.; Scott, D. 2006. Waterbird population estimates. Wetlands International, Wageningen, The Netherlands.

Further resources

IUCN SSC Cormorant Specialist Group

Search for photos and videos, and hear sounds of this species from the Macaulay Library

Media resources

Recommended citation
BirdLife International (2021) Species factsheet: Urile penicillatus. Downloaded from on 16/10/2021. Recommended citation for factsheets for more than one species: BirdLife International (2021) IUCN Red List for birds. Downloaded from on 16/10/2021.